Afton graptolites revisited

There are two great lost fossil sites in the Twin Cities area. (The Brickyards don't count; they're not lost, they just aren't open to collection.) One is the Johnson Street Quarry, where workers cut into a bed in the Hidden Falls Member of the Platteville Formation that had unusually abundant echinoderms. As described in Sloan et al. 1987: 200, "Sardeson mined out a spot in this unit in the old Johnson Street Quarry in Minneapolis (now filled with garbage, and covered with Interstate 35) that produced about 20 specimens of the starfish Protopalaeaster narrawayi, several specimens of the crinoid Cremacrinus arctus (Fig. 16.2), edrioasteroids, cystoids, brachiopods, bryozoans, molluscs, and graptolites." This is slightly out of date; instead of a dump, there's now a Quarry Shopping Center with a Cub Foods, Home Depot, and Target, although even with all those options you can't get an edrioasteroid there anymore. Regardless of the exact character of the overburden, it seems unlikely that anyone will be doing any paleontological follow-up there anytime soon. The other locality is the Afton graptolite locality in the St. Lawrence Formation. We already had a post on why this locality was important; what I'm curious about is where exactly it was. A locality, even if "lost", had to have been *somewhere*, and apart from the scientific and historic interest, there very well could be similar fossils in rocks nearby. Indeed, Hughes and Hesselbo (1997) reported graptolites in the lowest strata of the St. Lawrence Formation in their Afton section, where collection may have postdated the road work that destroyed the classic location. For some reason, despite its “classic” nature, nobody ever saw fit to just put a pin on the map. What clues do we have?

New page: Quaternary vertebrate inventories (USA)

There's an electronic stack of topics I've considered for blog posts, and within that a subset earmarked "do before hanging up the keyboard". One of these, almost from the very beginning, was a bibliography of Quaternary vertebrate locality inventories like the kind Oliver Perry Hay pulled together. Being a detail hoarder, I've always loved having this kind of information. Maybe I'm the only one, but it's my blog and if I want to put together a page of old locality lists, I'll do it. One might think "If I have to find this information, all I need is the Paleobiology Database" and call it a day, to which I just smile and nod politely. Anyway, you can find the page here.

Pop quiz! Mammoth or mastodon?

New thematic inventories and a bit more cave paleontology

A couple of thematic inventories have just been published with the involvement of the NPS Paleontology Program. One of these is the first part of a paleobotany series, eventually to be three parts covering the Cenozoic, Mesozoic, and Paleozoic. The first part, Matel et al. (2026), covers the Cenozoic and is available for download through February 24. This is one of the projects from the National Park Service-Paleontological Society Paleontology in the Park Fellowship Program. In it, we've worked to provide information on Cenozoic paleobotany for every park unit where it's known. (It's easier to do concisely when the record is one piece of petrified wood than when the park is, say, Florissant Fossil Beds National Monument, so necessarily the scale of detail varies.) For practical reasons the focus is on macrofossils, with pollen and such as an adjunct.

The other new inventory, Santucci et al. 2025, covers NPS cave paleontology from 2002 to 2023. This is presented as an update of a previous report (Santucci et al. 2001). I'm taking the opportunity here to add a postscript because the past couple of years have been very productive. First of all, we came upon a new park record: Kalaupapa National Historical Park, on the north side of Molokaʻi, Hawaii. Although main focus here is historical, this park has a very interesting geological story too, involving the collapse of a volcano and the subsequent growth of another smaller volcano. The smaller volcano is now extinct, leaving behind Kauhakō Crater. Inside the crater people have found bird bones in a cave. One is a partial ulna of the extinct flightless ibis Apteribis glenos, the other is a coracoid of Pterodroma hypoleuca, the modern Bonin petrel, which does not live on the island today (Olson and James 1982, 1991).

Also, a few things in press or under study have progressed in the past couple of years. Avid readers of this blog will know that the "musk ox" of Muskox Cave at Carlsbad Caverns National Park is now named Speleotherium logani (White et al. 2025). The Cumberland Bone Cave monograph has also been published (Eshelman et al. 2025). Most notable, though, has been the publication of work on Mammoth Cave National Park. The park inventory (Toomey et al. 2025) was mentioned here a few months back. There has also been a good short summary of the Mississippian vertebrates (Hodnett et al. 2024a) and no fewer than five new taxa: ctenacanthiforms Troglocladodus trimblei and Glikmanius careforum (Hodnett et al. 2023); petalodonts Clavusodens mcginnisi (Hodnett et al. 2024b) and Strigilodus tollesonae (Hodnett et al. 2024c); and holocephalan Macadens olsoni (Hodnett et al. 2025).

Clavusodens mcginnisi on the prowl, among the crinoids. Illustration by Benji Paysnoe for the National Park Service.

References

Eshelman, R. E., C. J. Bell, R. W. Graham, H. A. Semken, Jr., C. B. Withnell, S. G. Scarpetta, H. F. James, S. J. Godfrey, J. I. Mead, J.-P. Hodnett, and F. V. Grady. 2025. Middle Pleistocene Cumberland Bone Cave Local Fauna, Allegany County, Maryland: a systematic revision and paleoecological interpretation of the Irvingtonian, Middle Appalachians, USA. Smithsonian Contributions to Paleobiology 108.

Hodnett, J.-P., R. Toomey, H. C. Egli, G. Ward, J. R. Wood, R. Olson, K. Tolleson, J. S. Tweet, and V. L. Santucci. 2023. New ctenacanth sharks (Chondrichthyes; Elasmobranchii; Ctenacanthiformes) from the Middle to Late Mississippian of Kentucky and Alabama. Journal of Vertebrate Paleontology 43(3): e2292599. doi: 10.1080/02724634.2023.2292599.

Hodnett, J.-P., R. Toomey, R. Olson, K. Tolleson, R. Boldon, J. Wood, J. S. Tweet, and V. L. Santucci. 2024a. Sharks in the dark: Paleontological resource inventory reveals multiple successive Mississippian Subperiod cartilaginous fish (Chondrichthyes) assemblages within Mammoth Cave National Park, Kentucky. Park Stewardship Forum 40(1): 53–67. doi: 10.5070/P540162921.

Hodnett, J.-P. M., H. C. Egli, R. Toomey, R. Olson, K. Tolleson, R. Boldon, J. S. Tweet, and V. L. Santucci. 2024b. Obruchevodid petalodonts (Chondrichthyes, Petalodontiformes, Obruchevodidae) from the Middle Mississippian (Viséan) Joppa Member of the Ste. Genevieve Formation at Mammoth Cave National Park, Kentucky, U.S.A. Journal of Paleontology 98(6): 1087–1097. doi: 10.1017/jpa.2024.40.

Hodnett, J.-P., R. Toomey, R. Olson. J. S. Tweet, and V. L. Santucci. 2024c. Janassid petalodonts (Chondrichthyes, Petalodontiformes, Janassidae) from the middle Mississippian (Viséan) Ste. Genevieve Formation, Mammoth Cave National Park, Kentucky USA. Historical Biology 36(9):1783–1792. doi: 10.1080/08912963.2023.2231955.

Hodnett, J.-P., R. Toomey, H.-D. Sues, V. Santucci, K. Tolleson, and J. Tweet. 2025. A new euchondrocephalan chondrichthyan (Chondrichthyes, Euchondrocephali) from the Middle Mississippian (Viséan) Joppa Member of the Ste. Genevieve Formation at Mammoth Cave National Park, Kentucky, USA and a reassessment of the Lower Mississippian (Tournaisian-Viséan) “Helodus” coxanus Newberry, 1897. New Mexico Museum of Natural History and Science Bulletin 100: 87–93.

Matel, T. P., I. B. Huegele, C. R. Cace, K. M. M. Bober, L. D. Boucher, V. E. McCoy, E. J. Hermsen, S. R. Manchester, C. C. Visaggi, J. S. Tweet, and V. L. Santucci. 2026. Cenozoic paleobotanical resource inventory of the National Park System. Elements of Paleontology. Cambridge: Cambridge University Press. 10.1017/9781009770477.

Olson, S. L., and H. F. James. 1982. Prodromus of the fossil avifauna of the Hawaiian Islands. Smithsonian Contributions to Zoology 365: 1–59.

Olson, S. L. and H. F. James. 1991. Descriptions of thirty-two new species of birds from the Hawaiian Islands. Part I. Non-Passeriformes. Ornithological Monographs 45:1-88.

Santucci, V. L., J. Kenworthy, and R. Kerbo. 2001. An inventory of paleontological resources associated with National Park Service caves. National Park Service Geological Resources Division Technical Report NPS/NRGRD/GRDTR-01/02.

Santucci, V. L., J.-P. Hodnett, P. Seiser, J. S. Tweet, and J. Wood. 2025. National Park Service cave paleontology: 2002-2023. Journal of Cave and Karst Studies 87(4):108–116. doi: 10.4311/2024PA0119.

Toomey, R. S., J. S. Tweet, and V. L. Santucci , editors. 2025. Mammoth Cave National Park: Paleontological resource inventory (public version). Science Report NPS/SR—2025/243. National Park Service, Fort Collins, Colorado. https://doi.org/10.36967/2308547 

White, R. S., J. I. Mead, and G. S. Morgan. 2025. Logan's austral scrubox, a new ovibovine (Mammalia: Artiodactyla: Bovidae) from Muskox Cave, Eddy County, New Mexico. New Mexico Museum of Natural History and Science Bulletin 101: 473–494.

Your Friends The Titanosaurs: Yeneen houssayi

What do you get the formation that has five established titanosaur species and a couple of ne'er-do-wells? A sixth titanosaur, of course!

All joking aside, it's misleading to think of "Titanosauria" as a relatively small-scale group like Diplodocidae or Brachiosauridae when it's really a massive, sprawling complex including multiple distinct lineages that essentially monopolized all things sauropod in the Late Cretaceous. We talk of the Morrison Formation having diplodocids and dicraeosaurids and camarasaurids and brachiosaurids and whatever else, and that may sound more impressive than the Bajo de la Carpa Formation having six different titanosaurs. "What? You need all those titanosaurs?" "All those titanosaurs" are functionally replacing each of those smaller clades of the Morrison, as well as apparently just about everything else large and herbivorous. It's just the relationships and smaller divisions are still fuzzy. Eventually, we'll have a better grasp. But enough philosophy! Let's bring on today's guest, Yeneen houssayi. (And thank you to Alberta Claw and Stephen Poropat for supplying me with the paper!)

Silvisaurus condrayi

You wait long enough, and history starts to pile up, like falling snow (only history doesn't melt away and has a way of churning back to the surface). In paleontology, we've just passed two centuries with Megalosaurus (1824) and Iguanodon (1825). I was thinking about the topic of this post and how it was a relatively recent name, even though it felt old, but then realized that 1960 is not quite as recent as it was when I first read about Silvisaurus in the mid-1980s. I'm not tracking stats to any great degree, but it's probably safe to say there aren't a lot of people reading this who were mid-career when the scientific description of Silvisaurus condrayi was published.

Ghost of Christmas Presents

My first love in science, the first thing that really interested me intellectually, was astronomy. It was early 1986, around the time I turned five. (I was a weird kid.) Anyway, I was really into Star Wars, and wanted to go to space and explore different planets. I had (and still have) a book, "Astronomy Today", with lots of great illustrations and diagrams; sure, I wasn't *reading it* reading it yet, but I got a lot out of those illustrations. Then I discovered that, contrary to the movies, we couldn't exceed the speed of light, so we were basically stuck here. It's a nice enough solar system, but, well, y'know... the realization was terribly disappointing, and space went from an obsession to a pleasant side interest (I still love going to the planetary science sessions when attending GSA annual meetings).

At the time, we had just moved to Tucson. There's a lot of difference geologically between southern Arizona and south-central Minnesota; for starters, a lot more of the geology is right there at the surface in Arizona, and a lot more of it is Mesozoic. Still, Arizona is not Dinosaur Central; it's more like a nice suburb. After nearly 40 years I can't say what it was about dinosaurs that caught my eye. Maybe it was the British Museum toys at the Arizona-Sonora Desert Museum. Whatever it was, the feedback loop started: kid shows interest, people around start feeding the interest, interest intensifies...

The step that sealed the interest in paleontology came at Christmas 1986. There was one rectangular gift with unusual heft. Under the wrapping paper was the Normanpedia, "The Illustrated Encyclopedia of Dinosaurs". Could I read it? Well, okay, sure, I was a great reader for five-and-a-half, but it's safe to assume it took some time and practice before I was conversant (and the book probably accelerated the process, so on top of his scientific accolades, David Norman can take some credit for improving at least one kid's  literacy). But don't forget the second word of the title! The restorations, the photos, the skeletal diagrams! I went through reams of paper tracing skeletons, then drawing outlines around them on the other side of the sheets. That way, there would be a skeleton on one side, a drawing on the other, and if I held the paper to the light, both of them would be there, skeleton revealed inside the body. I loved that book to death; it's in a box of mementos somewhere at home, dust jacket long gone, spine held together by heavy-duty tape, a slip of paper taped over the "dinosauroid" drawing because it creeped me out. I had to get a second copy in the Nineties because the first was falling apart. The second copy also started falling apart after about ten years, so now I'm on my third. Anyway, no particular moral or commentary or insight this time, just wanted to share some holiday nostalgia. See you in 2026!

What do Delaware, Hawaii, and Rhode Island have in common?

Answer: They are the only three U.S. states for which I can't find a record of a mastodon or mammoth.

Hawaii is not exactly a surprise. Certainly proboscideans can swim, but if you want a large mammal that can swim halfway across the Pacific Ocean without dying, it's going to look and function suspiciously like a cetacean instead.

Delaware and Rhode Island, on the other hand, doubtless had mammoths and mastodons tromping over them at some point, because they're surrounded by states (and submerged coastal plains) that have their fossils. They're just both small states not blessed with the most helpful geology, and mantled by urbanization. Rhode Island gets an additional strike from being subjected to glaciers. Someday the fossils will turn up, though. Maybe there's already something; maybe somebody has a funny-looking doorstop they found in 1973 and never paid much attention to, or there's a notice in a local paper published in 1989 about a couple of campers who found a tooth and turned it over to a local historical society, or somebody will hop into the comments of this post and say, "Hey, ya numbskull, ya missed this!"

Now, the nitty-gritty: every once in a while I'd wondered how many states have records of mammoths or mastodons, and occasionally puttered in a source or two. This time I decided to be more thorough. I checked the Hay inventories, Neotoma (which has the old FaunMap contents), and the Paleobiology Database for starters. These three together narrowed down the list of absences to Delaware, Hawaii, New Hampshire, and Rhode Island. If you're curious: Hay didn't have mammoths or mastodons in those four plus Maine and basically omitted Alaska, but had all the rest plus the District of Columbia; Neotoma was missing the four plus Alabama, Mississippi, North Dakota, Vermont, and D.C.; and the PBDB was missing the four plus Connecticut, Louisiana, Massachusetts, Maine, New Jersey, North Dakota, Vermont, and D.C. This is why it's important to check multiple sources; no one source has everything.

Putting aside Hawaii, that left Delaware, New Hampshire, and Rhode Island to check out. Now was the time to drill into the resources of various search engines and museum databases/iDigBio. For the record, I leaned on Google Scholar, Google Books, and the news tab of regular Google (not Google News), with various permutations of "mammoth", "mastodon", "Mammuthus", and "Mammut" plus a state, with additional terms as necessary to knock out particularly obnoxious false positives. This eliminated New Hampshire, which now has a record from a find on land after a couple of teeth found nearby offshore. For a short time I'd thought I'd found one for Delaware, thanks to a short piece in the Winter 1994 issue of "First State Geology" (p. 3). The article mentioned that Jeremy Cloutier of Milford had donated a mastodon tooth to the Delaware Geological Survey, but the tooth derived from offshore clam dredging and therefore doesn't count for our purposes. So, for now, we're still at Delaware, Hawaii, and Rhode Island. (Still a few field days 'til Christmas, if you're feeling like pulling off a Christmas miracle!)

"Archie", the world's largest mounted Columbian mammoth skeleton, on display in Elephant Hall, University of Nebraska State Museum.